Chuck Darling and Kelly Carr are reunited as lead news anchors ten years after their split. Can the duo rekindle the on-screen magic they once had? Or will history repeat itself once again?
Chuck is given a goldfish as a gift, which makes him feel like he needs to prove himself as a good caretaker to Kelly. To do so, Chuck has Ryan help him keep up the illusion that the goldfish is happy, healthy and still living. Meanwhile, while the search for a gripping news story is in full swing, Crezyzewski prepares for a hands-on police taser story.
Chuck and Kelly continue to get on each other's nerves both on and off camera. However, things change when Chuck's dinner date cancels and he runs into Kelly and Gracie having dinner, which is the making for an awkward meal when he joins them. Gracie and Chuck hit it off, and despite Kelly's disdain for Chuck, she begins to think he might be good for Gracie. Back at the station, Gary is helped by Marsh to make his first anchor highlight video, but it doesn't turn out how they hoped. Meanwhile, Ryan tries to hide his obsession with Montana.
After running into some old news buddies, Chuck feels that he must prove he still has what it takes to be a network anchor. To do so, Chuck tries to trick Kelly so he can cover the lead story of the night, but when the lead story switches back and forth, Kelly and Chuck try to convince each other as to which story each one should cover.
Chuck turns down a dinner invitation from Ryan, leaving his Friday night wide open for romance. When Kelly tells Ryan he's being selfish, Chuck accuses Kelly of being too uptight. Each wants to prove the other wrong: Chuck agrees to dinner with Ryan and Kelly heads out with Montana for a night on the town.
Gracie spends time at the station to stay clear from a bully, while Chuck and Kelly disagree on how to handle the situation. Meanwhile, Gary and Marsh bet on baseball; Ryan has a lot in common with one of Gracie's classmates; and a little girl takes Montana's forecast by storm.
A trapped raccoon in the attic disrupts Kelly's plans for Gracie's birthday party. The whole crew attempts to capture the critter before Gracie arrives home.
Chuck agrees to babysit Gracie when Kelly is asked to deliver a eulogy for an anchor, who unbeknown to her, had been telling everyone that he and Kelly were secret lovers.
Chuck becomes jealous when Kelly is asked out by a younger, handsome rival. Meanwhile, Gary chastises the members of an animal costume convention.
Kelly and Chuck discover an incriminating photo from their past that they want destroyed, while attending a surprise party in which Marsh has made for Chuck. Meanwhile, Gary and Ryan make a shocking discovery of their own while in Marsh's bedroom.
Kelly gets cold feet as she and Chuck were about to reveal to Gracie that Chuck is her real father. She decides that they need to resolve a few issues first and suggests they visit a therapist, which only complicates matters more when it's discovered that they both are holding back pent-up frustrations. Meanwhile, Gary is upset when he fails to receive the hug and words of encouragement which the security guard hands out daily.
As Gracie deals with the realization that Chuck is her father, Chuck and Kelly remember the night that changed their lives forever.
When newly appointed WURG station manager Ginger Ko decides to make some unpopular changes to the newsroom, Chuck and Kelly pay her a visit. Despite their intentions to confront her about the proposed changes, Ginger turns on the charm, sending Chuck and Kelly away feeling good about the transition. However, the aftermath from their seemingly innocent exchange causes quite a stir in the newsroom, especially for Ryan, who has to step up and carry out a career-first responsibility. Meanwhile, Gary is apprehensive about booking a cruise with his wife for fear the new boss won't agree to him taking the time off, and Marsh accidentally takes his wife's estrogen pill.
After deciding the best way to disclose their parenthood, an untimely ad campaign from the station changes their plans. Meanwhile, Gary begins to regret Marsh's generosity after receiving a gift from him.