The Beast Wars on prehistoric Earth are over and the Maximals have won and preserved the past for the battle between the Autobots and Decepticons. The Maximals have used the shuttle from the Ark and are returning to Cybertron. Upon arriving, the crew blacks-out and awakes to a desolate and barren home with no fellow transformers anywhere. Furthermore, many of the returning Maximals are missing as well. Only Optimus Primal, Cheetor, Rattrap, and Blackarachnia awake together. They are attacked and outnumbered by drones and are forced into hiding. They must now find their friends and be willing to go through a change that will turn their sparks upside-down.
Optimus is alone. All of the other Maximals have had their sparks removed. Now it is the final battle between Optimus Primal and Megatron in his new, extremely powerful form. Can Optimus finally defeat Megatron? Or, will Megatron finally make the Cybertron that he has envisioned for so long?