The series begins in 52 BC, as Gaius Julius Caesar completes his conquest of Gaul after eight years of war, and prepares to return with his army to Rome. While Caesar's self-interested niece Atia and long-lost paramour Servilia anxiously await the general's return, ruling patricians despair that Caesar's homecoming will disrupt the status quo, and threaten the extravagant prosperity they've enjoyed at the expense of the lower class. In the Senate, old-guard leaders plot to undermine Caesar's influence by convincing his old friend, Pompey Magnus, that the general is a threat. Back at the front, two mismatched soldiers, Lucius Vorenus and Titus Pullo, see their fortunes rise following several serendipitous missions. As Caesars legions move closer to Rome, allegiances are put to the test for soldiers and civilians - and the escalating tensions climax with a full-scale conflict destined to change history.