In the superficial world of high fashion, image is everything. Styles come and go, and the only constants are the superthin beauties who wear them. How can an ordinary girl a slightly plump plain-Jane from Queens possibly fit in?
Betty unwittingly loses the book, a proof of the entire magazine, complete with un-retouched photos of a popular actress who looks less than lovely without a little computer magic. If the photos get out, it could create huge embarrassment for Mode. Wilhelmina plots to use the snafu to bring Daniel down. Meanwhile, Bradford Meade secretively inquires about Fey Sommers final effects.
Betty uses her Queens connections to lure big time photographer (and former Queens boy) Vincent Bianchi to shoot for Mode. But when Bianchi insists that Betty meet at a posh restaurant, she feels pressured to get a makeover only things dont quite turn out as planned. Meanwhile Wilhelmina primps for a big date, and Bradford tries to get his hands on Fey Sommers death car.
When Mode is forced to abandon its big Christmas photo shoot because another magazine has copied the idea, Betty forms an uneasy alliance with Marc and Amanda. The three fear they may have leaked the idea to someone at the rival mag during happy hour . . . and that their jobs will be history if anyone finds out. Meanwhile, Daniel and Wilhelmina reluctantly work together to come up with a new photo spread as Mode's deadline looms, and Betty's nephew, Justin, is oblivious to the turmoil as he spends a day at the magazine for a class project.
It's Halloween and Betty is in for a series of frightful experiences, ranging from showing up dressed for the Mode costume contest . . . only to learn there isnt one, to dealing with her growing attraction to Henry -- while a jealous Walter fumes -- and, worst of all, tracking down a watch Daniel left at one of his many girlfriends apartments. Meanwhile, Daniel pays a visit to his mom in rehab, and Wilhelmina is horrified to learn that she can't fit into a designer gown.
Daniel falls in instant lust with a mysterious beauty he sees in the Mode lobby (guest star Salma Hayek) and asks for Bettys help in finding out the ladys identity. Meanwhile, Betty and Hilda scramble to raise cash when they get some unexpected news, and Wilhelmina must deal with an unexpected house guest, her daughter.
Betty takes a break from her escalating family troubles when she's assigned to review a posh new hotel. Meanwhile, Daniel is dismayed when Sofia Reyes and her team take up residence in the Mode conference room, and Wilhelmina goes a little bit country to woo a mega rich Texan to advertise his stores in Mode.
Its Thanksgiving Day and everyone at Mode celebrates in their own special way. Betty is torn between work and home; Daniel attends his parents brunch in hopes of seeing Sofia; Marc and Amanda camp out at the Mode offices, and Wilhelmina attempts something shes never done before cooking a turkey.
The day after Thanksgiving is no holiday for Betty, who struggles to pull together a last-minute Mode photo shoot with a famous couple and their baby, while a dejected Daniel recuperates from a hangover at the Suarez house.
Amanda hopes her successful holiday office party will be the gift that keeps on giving. Meanwhile, Betty sorts through her feelings for Walter and Henry; Daniel's love for Sofia grows; and Marc tries to stay on Wilhelmina's good side.
Christina's swag lottery creates a feeding frenzy among the office fashionistas, while Betty wears herself out helping a suddenly cash-poor Daniel entertain a famous Japanese designer.
Daniel's future plans with Sofia are thrown into question when Betty makes a startling discovery; Wilhelmina's romance with Ted changes her mood at the office; and Ignacio doesn't see eye-to-eye with his immigration caseworker.
Daniel is suffering a broken heart and Betty decides to help him get over Sofia by setting him up with a super model. Meanwhile Wilhelmina is picking people to fire or keep when she becomes editor-in-chief.
During Fashion Week in New York City, Betty works to make sure Mode's runway show goes flawlessly. However, Daniel has Hilda help her with the show, much to Betty's dismay. Marc and Amanda continue on a mission to seal the season's hottest fashion item; Daniel is surprised when his father decides to hand over Meade Publications, but a visit from an old friend sends him back to his old ways; and the Mode fashion show is suddenly stopped by a shocking revelation.
The masked woman makes an announcement that leaves Mode in disarray. Meanwhile, Betty tries to moderate the chaos, but Wilhelmina continues to plot her next move; Henry asks out Betty; and Justin is sent home from school for fighting.
Daniel is forced to reconnect with Grace Chin, a girl he once ditched on a date in college but is now New York City's most powerful lawyer. Betty meets a new friend, Charlie, in the Mode cafeteria, but soon learns she is Henry's girlfriend. Meanwhile, Christina and Marc are trapped together during a blizzard and work on a gown for Sarah Jessica Parker; Wilhelmina and Alexis get stranded at a bar; and Santos and Justin get stuck on the subway.
Betty asks her orthodontist, Dr. Farkas, to be her date to Charlie's birthday party after being invited by Henry. Meanwhile, Wilhelmina tries to tear Daniel and Alexis apart when she learns that Daniel was the one who hired Grace Chin to defend their father; and Amanda piques the interest of an unusual designer.
Marc convinces Betty to pretend to be his girlfriend after his mother, Mrs. Weiner, arrives in town. However, when Mrs. Weiner invites herself to dinner at the Suarez home, Betty's family is forced to play along. Meanwhile, Daniel and Alexis have difficulty working together when their mother names them both co-editors-in-chief.
Betty tries to keep Daniel focused on work after he begins to slip back into his partying habits. Things get worse when a writer gets a hold of Fey Sommer's diaries. Meanwhile, Ignacio is pressured by the immigration caseworker to get into a romantic relationship with her; and Wilhelmina and Alexis try to get a hold of Fey's diaries, but each have a different reason for wanting to do so.
Having been disappointed by Daniel, Christina and Henry, Betty does her best to avoid them. Meanwhile, love is all around at Mode with Alexis being asked out on her first date as a woman; Amanda flirts with a man she assumed was gay; Wilhelmina is tired of dealing with her latest conquest; and Hilda is surprised by Santos. Also, Daniel is on his own dealing with his one time date with Petra.
Betty tries to find the cash to fix the latest Suarez family problem, all while working on the plans for Mode's assistant week bash. Meanwhile, Daniel mulls over Betty's advice on his sexual addiction; Alexis debates Rodrigo's offer; Hilda takes her cosmetology school exam; and Wilhelmina concocts a plan to take over Mode.
Betty and the Suarez family head to Guadalajara to fix Ignacio's immigration problems. Meanwhile, Jordan visits Alexis; Marc is jealous when Amanda finds a new best gay friend in Tavares; and Wilhelmina gives her new man a makeover.
Betty and Henry's relationship heats up as Wilhelmina makes a critical move in her power play attempt for Mode. Meanwhile, the Suarez's trip to Mexico takes a deadly turn.
A series of unfortunate events puts Betty's life in chaos, but she continues to deny how much she misses Henry. Meanwhile, Amanda finds comfort in junk food and Marc after learning that Fey Sommers is her biological mother; and Wilhelmina plots to use the Meade family events to her benefit.
Betty learns that Wilhelmina's new bodyguard is taking his job to another level and has even more weighing on her mind when Henry returns. Meanwhile, Bradford uses Alexis' amnesia to his advantage in order to rewrite family history; Hilda deals with recent events; and Amanda is given an inheritance from Fey Sommers.
With Henry back, Betty becomes distracted by another guy, Gio, who happens to be Mode's sandwich guy. As the people at Mode prepare for the social event of the season -- the Meades' Black and White Ball -- Wilhelmina tries to get her wedding back on track. Meanwhile, Amanda prepares for her first appearance as Fey Sommer's daughter and Claire invites herself to the party.
Betty decides to become a writer and takes a class, but soon finds it too challenging and ends up submitting a Mode story instead of her own. Alexis, who is still suffering from amnesia, returns to work with no memory of how to act like a girl. Meanwhile, a top advertiser tells Daniel to fire his sister and Amanda uncovers a secret about Wilhelmina.
Betty decides to give Internet dating a try and soon gets asked out on a bowling date, much to Henry's dismay. Meanwhile, Marc is attracted to someone he normally wouldn't be; Daniel and Alexis prep for a "family" dinner with Wilhelmina; and Hilda is unaware of Justin's ditching school.
Betty and Henry continue to see each other and plan a date to see "Wicked," but things don't go as they had planned when Daniel, thinking she's dating Gio, gives him the tickets. Meanwhile, Daniel tries to seduce an advertiser with the reputation of dating younger men.
While Wilhelmina prepare for her big day, things are thown into a spin when she learns that her maid of honor, Victoria Beckham, is getting more attention than she is. Meanwhile, Betty feels guilty over her pact with Wilhelmina to not tell Daniel about his future step-mother's infidelity with her bodyguard. Ignacio prepares for becoming a U.S. citizen, but when Betty moves in with Henry, his mind wanders elsewhere.
Mode's weekend receptionist helps Henry work overtime to get Betty something she needs. Wilhelmenia's romantic saga with Bradford needs new direction.
Everyone at Mode scrambles to fix the issue before the deadline. Daniel and Betty go to all lengths necessary to get the issue's cover girl out of rehab for a re-shoot. Alexis must charm the printer into giving them an extended due date.
Wilhelmina hits a snag in her newest image makeover plan. Meanwhile, Betty drags a reluctant Henry as her date along with Hilda and Gio.
Amanda finds a new way to search for her father's true identity. Meanwhile, Betty convinces Daniel to change Mode's message; and Wilhelmina receives unexpected news.
Henry and Betty are both effected weirdly by her new perfume, which was designed by the late Fey Sommers. A trail begings for Clair, accused of killing Fey, while Wilhelmina continues her hopes to become part of the Meade family.
Betty is assigned to interview a writer, but her assignment goes awry during a case of mistaken identity. Meanwhile, Daniel's new love is getting closer to him than he is aware of; and Marc helps Amanda with the search for her father.
Plans for a romantic 24th birthday alone with Henry are put on hold when Charlie stops by without warning. Meanwhile, Wilhelmina tries to ruin Renee's relationship with Daniel by hinting at dark secrets Renee is keeping.
Renee, who is becoming increasingly delusional, thinks that Betty is in love with Daniel, leading her to compete against her for his attention. Meanwhile, Hilda's new clinet turns out to be nemesis Gina Gambarro, who has returned to flaunt her wealthy husband in front of Hilda.
Betty tries to befriend Charlie, her rival, by throwing her a baby shower which turns out to be a party no one will forget. Meanwhile, Hilda talks to Justin's PE teacher when she finds out Justin is failing gym class. Elsewhere, Daniel tries to come to grips with his break-up with Renee; and Christina learns who is the father of the baby she's carrying for Wilhelmina.
Betty fights her attraction towards Gio, while still missing Henry, but find themselves together after agreeing to chaperon Justin's school dance. Meanwhile, Hilda does what she can to get Coach Diaz to notice her; Amanda agrees to do a reality show with her father, Kiss rock legend Gene Simmons; and Wilhelmina's return to Mode had Daniel feeling the heat amid a media frenzy.
Gio makes Betty an offer, but Henry also comes back to town with an offer of his own for Betty. Meanwhile, Betty plays center field for the charity Mode vs. ELLE Magazine softball game where Wilhelmina vies to strike out ELLE's star -- supermodel Naomi Campbell.
Betty must make a life-altering decision about what to do with her life. Meanwhile, Ignacio lands a job at a fast food restaurant while Wilhelmina forces Daniel to make changes in his life.
Wilhelmina, getting ready for her first issue as editor, offers Betty a job as her assistant. Meanwhile, Daniel learns he will have to undergo a custody battle if he wants to keep Daniel Jr. in the U.S. Later, Hilda's romance with Coach Diaz continues to heat up.
A tragic accident leaves the Mode staff stunned and Betty must find out what really happened when a detective comes looking for answers. Meanwhile, Ignacio learns about Hilda and Coach Diaz.
Gio returns to town, forcing Betty to attempt to make amends. However, Gio resists her attempts to make things right. Meanwhile, Daniel must cope with shocking family news.
Kimmie Keegan goes to Betty for help in finding a job. Meanwhile, Daniel decides to try a matchmaking service and gets a shocking "perfect" match. Elsewhere, Justin tries out for a Broadway musical.
The newest citizen Ignacio is ready to vote for the first time. Kimmie's true colors show as she competes against Betty. Meanwhile, with a large crowd passing through for the voting, Hilda uses the time to promote her shop and Wilhelmina prepares to hire a new financial officer whom Daniel hates.
Betty wants to be noticed by her cute neighbor, Jesse, by coming home later each night. Meanwhile, Amanda unexpectedly moves in with Betty after having nowhere else to go. Elsewhere, Marc and Cliff mull over the idea of moving in together.
It will mean a nightmare for the PR department if Betty doesn't get a hold of Daniel before the next issue of Mode is released, but he is away at a company retreat. Meanwhile, Hilda finds out upsetting news about her styling salon business permit and Conner's fiancee meets the staff of the Mode.
Betty challenges her rival, Marc, for the final place in YETI, the Young Editors Training Program, which is a real competitive program. Meanwhile, it seems that the others are each attracted to another...Daniel is after Molly, Wilhemina is after Connors and Justin makes for an unlikely friend.
Betty and Amanda are cash strapped and decide to use this to their benefit while spending a night on the town by writing an article on Mode's website named "How I Blew Ten Grand Without Spending a Dime." The girls receive an evening in New York City which they will never forget.
Betty has to juggle work and family as she tries to impress Jodie Papadakis, the editor for YETI. The rivialy heats up after an encounter with Teri, an assistant at Elle Magazine.
Betty is torn between work and home following a family crisis. Meanwhile, couples Daniel and Molly and Connor and Wilhelmina try to hide their relationships.
Betty must decide whether or not to make a personal decision in order to help her family's money problems, but is sidetracked by her sexy neighbor Jesse. Meanwhile, Daniel tries to cover his tracks when Connor works to find out who is Molly's mystery man.
Betty makes a mistake while preparing a special video for Claire's 60th birthday, when the spotlight is accidentally shined on Ignacio. Meanwhile, Daniel and Wilhelmina's relationship is facing similar obstacles.
On assignment from YETI, Betty makes a shocking discovery in regards to Mode's yearly budget. Meanwhile, Betty has to deal with more attention from Matt as well as unexpected news from Henry.
The Suarez family have to come up with enough money to buy their home after their landlord decides to sell it. Meanwhile, the Mode staff copes with recent events.
Betty meets Matt's mother, but the meeting turns into a disaster. Meanwhile, Wilhelmina comes up with a way to solve the company's cash problem; Daniel deals with his relationship with Molly; and the Suarez's work together to set Hilda up with Archie.
Betty receives a makeover from Marc and Amanda in hopes of taking her romance with Matt in a different direction. To pump up the bottom line for the Mode's next issue, Daniel and Wilhelmina try adding some spice into it. Hilda ponders her relationship with Archie.
The Suarez family meets Matt's billionaire dad.
Baby William iѕ kidnapped - аnԁ Betty iѕ dragged іnto thе ensuing drama. meanwhіlе Archie (guest star Ralph Macchio) asks Hilda tо bе part оf hіѕ latest political campaign.
Betty teams wіth Matt аnԁ Marc fоr thеіr all-important final YETI project.
Just aѕ Betty agrees tо move іn wіth Matt, Henry pays а visit tо NYC, forcing Betty tо admit ѕhе ѕtill hаs feelings fоr hеr fоrmеr fiance.
Betty аnԁ Marc compete fоr thе ѕame job.
Betty finds that everything is different as she starts her new job as associate features editor. Matters are made worse when she finds that her boss and ex-boyfriend, Matt Hartley, is still upset over their breakup. Meanwhile, without Betty, Daniel begins to spiral downhill and continues mourning the loss of his wife; Justin starts high school, but hides the fact that bullies are taunting him; and Wilhelmina continues to harbor a deep secret.
Betty gets the opportunity to run a Mode photo shoot, which is for the United Nation's anti-malaria campaign, "Nothing but Nets." Meanwhile, Amanda bonds with Daniel's temp, Helen, and Wilhelmina does everything possible to protect a loved one.
Marc is appointed as Betty's temporary assistant by Wilhelmina. Betty decides to do a story on the worst jobs in New York. Meanwhile, Hilda causes trouble for Archie's campaign and Daniel hires himself a beautiful but inept assistant.
Ignacio puts together a "multi-culti" holiday dinner for Jean, who is Jewish, while Betty and Hilda both have pregnancy scares. Both Betty and Hilda decide to buy Be-Shure instant pregnancy tests from a pharmacy where Jean and Ignacio happen to be shopping. Meanwhile, Hartley finds a replacement for Wilhelmina -- Denise Ludwig, a clueless mogul from Hollywood -- but Marc puts together a plan to get Wilhelmina back in control.
Worried she's spending too much time with Matt, Betty tries to find something else for Matt to be involved in, which turns out to be a gallery showing of his art work. However, the exhibit creates some unexpected results. Meanwhile, Marc learns his one-night stand is falling for him; Hilda debates telling Bobby that he's the father of her child; and Hartley hires a replacement for Wilhelmina.
As flashbacks tell the story of the last few days, Betty talks to Mr. Z about her worst week ever. However, when Wilhelmina has Betty work on various experiments, the new tasks nearly kill her. Meanwhile, Daniel learns that Marc is trying to sabotage him, and Wilhelmina makes a commitment to Connor.
Betty's building is robbed right before the karaoke party, and Betty is partly to blame for the break-in. Meanwhile, Amanda tricks Daniel and Wilhelmina into getting counseling; and Ignacio refuses to accept that Bobby is now part of Hilda's life.
Wilhelmina is furious when she learns about Wilhediva Hater, her drag queen impersonator. Meanwhile, Betty's blog wins her an award, which creates tension between her and Daniel; Amanda rehires Helen, but Wilhelmina later orders Amanda to fire her; and Marc runs his own photo shoot.
Wilhemina marginalizes Betty at Fashion Week, but Daniel has her reassigned to cover a low-priority show. There, Betty meets Marisa, one of the "10 Designers to Watch." Meanwhile, Amanda teams up with Helen to start a new line, but both Marc and Betty refuse to show their work to Wilhelmina.
To speed along the investigation into her house fire, Betty is pressured to date an annoying fireman, Jimmy. Wilhemina tries to prove to Don, an old flame from her past, that she's just "Wanda from the block" instead of an uber-bitch, and Amanda draws closer to Tyler, convincing Claire to hire him as an in-house model at Mode. Meanwhile, Hilda meets Bobby's parents for the first time.
Justin loves his acting class, helmed by the eccentric Lena Korvinka, and finds himself in an unexpected triangle with his new pals, Austin and Lily. When Betty drops by class, she's smitten with a cool young playwright, Zachary, whom she will profile for Mode. But she challenges her own ethics when she starts dating him. And Wilhemina is hospitalized with a perforated ulcer, and can't help her type-A personality even when bedridden, but then her roommate, Roberta, prompts an epiphany.
After being knocked unconscious at a photo shoot, Betty has a dream in which her orthodontist shows her how her life would have been had she never worn braces.
Amanda and Hilda join Betty on a trip to Fashion Week in London, where Betty reconnects with Christina (Ashley Jensen) and has a chance encounter with Gio (Freddy Rodriguez). Meanwhile, Wilhelmina goes to extremes to undermine Claire and Daniel by getting close to Tyler, and Bobby wonders why Justin and a classmate have become inseparable.
Betty asks Daniel to go to Hilda's wedding with her; Amanda wants to fix Marc up with her new client; Betty considers unexpected offers; Wilhemina uses Tyler as a pawn to fight the Meade empire.